Education:

B.S. Biological Sciences, California State Polytechnic University , Pomona , 1976.

M.S. Biological Sciences, California State Polytechnic University , Pomona , 1978.

Ph.D. Biological Sciences/Population Genetics, The University of California , Riverside , January 1985.

Postdoctoral Research, 1985, The Museum of Natural History , Los Angeles , California . Under the direction of Dr. John Wright, Curator of Herpetology.

Postdoctoral Research, 1986, University of California , Riverside . Under the direction of Dr. Michael Woodburne.